{"id":164,"date":"2020-06-22T09:53:02","date_gmt":"2020-06-22T16:53:02","guid":{"rendered":"https:\/\/wordpress.viu.ca\/biol223\/?page_id=164"},"modified":"2021-09-14T10:21:42","modified_gmt":"2021-09-14T17:21:42","slug":"arbutus-menziesii","status":"publish","type":"page","link":"https:\/\/wordpress.viu.ca\/biol223\/arbutus-menziesii\/","title":{"rendered":"Arbutus menziesii"},"content":{"rendered":"\n<p>Family: Ericaceae<\/p>\n\n\n\n<p>Common name: Arbutus<\/p>\n\n\n\n<p>E-flora BC: <a href=\"http:\/\/linnet.geog.ubc.ca\/Atlas\/Atlas.aspx?sciname=Arbutus%20menziesii\">http:\/\/linnet.geog.ubc.ca\/Atlas\/Atlas.aspx?sciname=Arbutus%20menziesii<\/a><\/p>\n\n\n\n<p>Wikipedia: <a href=\"https:\/\/en.wikipedia.org\/wiki\/Arbutus_menziesii\">https:\/\/en.wikipedia.org\/wiki\/Arbutus_menziesii<\/a><\/p>\n\n\n\n<p>Arbutus is a distinctive tree in our region, it is hard not to notice it with its smooth rusty-colored trunks and shiny green leaves. Very drought tolerant, it inhabits exposed rocky sites along the coast line, or rocky outcrops within a forest. It is our only evergreen broad-leafed tree, keeping its leaves on year round. Leaves that have become old and shaded by new growth are often shed in the summer. <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"1024\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-1024x1024.jpg\" alt=\"\" class=\"wp-image-151\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-1024x1024.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-300x300.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-150x150.jpg 150w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-768x768.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-1536x1536.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy-788x788.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/large-arbutus-copy.jpg 1920w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Arbutus in its typical habitat &#8211; a rocky outcrop.  Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<p>The bark of arbutus is one of its most distinctive features. The outer layer of the bark peels away in thin sheets every spring, revealing a smooth, initially green surface. Later in the summer, the green bark takes on a rusty-red color. <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy-1024x683.jpg\" alt=\"\" class=\"wp-image-141\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy-1024x683.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy-1536x1024.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy-788x525.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark-3-copy.jpg 1920w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>An arbutus stem in spring. The outer layer of bark peels off, revealing smooth green bark underneath. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-1024x683.jpg\" alt=\"\" class=\"wp-image-150\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-1024x683.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-1536x1024.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-2048x1365.jpg 2048w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-stem-7230-788x525.jpg 788w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>By summer, the green bark has taken on a rusty-red color. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"683\" height=\"1024\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-683x1024.jpg\" alt=\"\" class=\"wp-image-143\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-683x1024.jpg 683w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-200x300.jpg 200w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-768x1152.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-1024x1536.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-1365x2048.jpg 1365w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-788x1182.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-bark2-copy-scaled.jpg 1707w\" sizes=\"auto, (max-width: 683px) 100vw, 683px\" \/><figcaption>Very old portions of the trunk, at the base of the tree, accumulate layers of bark and take on a dark-reddish brown cracked appearance. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"768\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy-1024x768.jpg\" alt=\"\" class=\"wp-image-144\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy-1024x768.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy-300x225.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy-768x576.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy-1536x1152.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy-788x591.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-branching-copy.jpg 1920w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Why do arbutus trees shed their bark? It&#8217;s anyone&#8217;s guess. It is likely that the green bark has photosynthetic capacity in the spring. Compare the accumulation of lichens on the dead part of this branch, to the live (still shedding every spring) part. Perhaps shedding bark keeps trunks free from lichens and other debris. Whether this is adaptive or not is unclear. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<p>Arbutus leaves are firm and leathery, the tree invests a lot of matter and energy in constructing them, they will last for a while. Leaves of mature trees have smooth edges, while seedling leaves have fine toothed edges &#8211; this can make identification of seedlings a bit confusing. The leaves are arranged in an alternate \/ spiral patterns along the stem. <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy-1024x683.jpg\" alt=\"\" class=\"wp-image-146\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy-1024x683.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy-1536x1024.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy-788x525.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-leaves-copy.jpg 1920w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Egg shaped shiny leaves of arbutus. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"768\" height=\"1024\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy-768x1024.jpg\" alt=\"\" class=\"wp-image-147\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy-768x1024.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy-225x300.jpg 225w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy-1152x1536.jpg 1152w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy-1536x2048.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy-788x1051.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-leaves_leaf-miner-copy.jpg 1920w\" sizes=\"auto, (max-width: 768px) 100vw, 768px\" \/><figcaption>You&#8217;ll often notice the meandering traces of the arbutus leafminer (<em><a rel=\"noreferrer noopener\" href=\"https:\/\/pnwhandbooks.org\/node\/6950\/print\" target=\"_blank\">Marmara a<\/a><\/em><em><a rel=\"noreferrer noopener\" href=\"https:\/\/pnwhandbooks.org\/node\/6950\/print\" target=\"_blank\">rbutiella<\/a><\/em>) on arbutus leaves. The larvae of this insect feed on the tissue in the center of the leaf (the mesophyll), while staying protected from predators by the tough outer layer (the epidermis) of the leaf. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<p>Arbutus bears large clusters of white, somewhat fragrant, flowers in spring.<\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"799\" height=\"502\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-flower_Lotus-Johnson.jpg\" alt=\"\" class=\"wp-image-171\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-flower_Lotus-Johnson.jpg 799w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-flower_Lotus-Johnson-300x188.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-flower_Lotus-Johnson-768x483.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Arbutus-flower_Lotus-Johnson-788x495.jpg 788w\" sizes=\"auto, (max-width: 799px) 100vw, 799px\" \/><figcaption>Cluster of arbutus flowers in spring. Photo credit: Lotus Johnson<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy-1024x683.jpg\" alt=\"\" class=\"wp-image-145\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy-1024x683.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy-1536x1024.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy-788x525.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-flower-copy.jpg 1920w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Compare the urn-shaped arbutus flowers to those of <a rel=\"noreferrer noopener\" href=\"https:\/\/wordpress.viu.ca\/biol223\/gaultheria-shallon\/\" target=\"_blank\">salal<\/a> and red huckleberry, and you&#8217;ll note the kinship among them &#8211; they are all members of the Ericaceae family, where many members have this flower shape. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<p>After pollination, the flowers produce striking clusters of berries. Depending on the tree, these range from bright orange to deep red in colour. <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy-1024x683.jpg\" alt=\"\" class=\"wp-image-148\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy-1024x683.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy-1536x1024.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy-788x525.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/arbutus-over-water-copy.jpg 1920w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Flowers pollinated in spring develop into clusters of orange berries that appear in early fall. Photo credit: Douglas Fraser<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"800\" height=\"600\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_cynthia-cheney.jpg\" alt=\"\" class=\"wp-image-170\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_cynthia-cheney.jpg 800w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_cynthia-cheney-300x225.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_cynthia-cheney-768x576.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_cynthia-cheney-788x591.jpg 788w\" sizes=\"auto, (max-width: 800px) 100vw, 800px\" \/><figcaption>Cluster of bright orange-red arbutus berries in fall. Photo credit: Cynthia Cheney<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"591\" height=\"600\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_ME-Sanseverino.jpg\" alt=\"\" class=\"wp-image-169\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_ME-Sanseverino.jpg 591w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/berries_ME-Sanseverino-296x300.jpg 296w\" sizes=\"auto, (max-width: 591px) 100vw, 591px\" \/><figcaption>Close-up of the leathery skin of arbutus berries. The berries are fleshy, but taste very astringent due to high tannic acid content. They do not appear to have been used much for food by First Nations people of our area. Photo credit: ME Sanseverino<\/figcaption><\/figure>\n\n\n\n<p><\/p>\n","protected":false},"excerpt":{"rendered":"<p>Family: Ericaceae Common name: Arbutus E-flora BC: http:\/\/linnet.geog.ubc.ca\/Atlas\/Atlas.aspx?sciname=Arbutus%20menziesii Wikipedia: https:\/\/en.wikipedia.org\/wiki\/Arbutus_menziesii Arbutus is a distinctive tree in our region, it is hard not to notice it with its smooth rusty-colored trunks [&hellip;]<\/p>\n","protected":false},"author":2649,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_monsterinsights_skip_tracking":false,"_monsterinsights_sitenote_active":false,"_monsterinsights_sitenote_note":"","_monsterinsights_sitenote_category":0,"footnotes":""},"class_list":["post-164","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages\/164","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/users\/2649"}],"replies":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/comments?post=164"}],"version-history":[{"count":10,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages\/164\/revisions"}],"predecessor-version":[{"id":970,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages\/164\/revisions\/970"}],"wp:attachment":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/media?parent=164"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}