{"id":252,"date":"2020-06-24T13:30:49","date_gmt":"2020-06-24T20:30:49","guid":{"rendered":"https:\/\/wordpress.viu.ca\/biol223\/?page_id=252"},"modified":"2020-07-07T16:57:54","modified_gmt":"2020-07-07T23:57:54","slug":"alnus-rubra","status":"publish","type":"page","link":"https:\/\/wordpress.viu.ca\/biol223\/alnus-rubra\/","title":{"rendered":"Alnus rubra"},"content":{"rendered":"\n<p>Family: Betulaceae<\/p>\n\n\n\n<p>Common name: Red alder<\/p>\n\n\n\n<p>E-flora BC: <a href=\"https:\/\/linnet.geog.ubc.ca\/Atlas\/Atlas.aspx?sciname=Alnus%20rubra\">https:\/\/linnet.geog.ubc.ca\/Atlas\/Atlas.aspx?sciname=Alnus%20rubra<\/a><\/p>\n\n\n\n<p>Wikipedia: <a href=\"https:\/\/en.wikipedia.org\/wiki\/Alnus_rubra\">https:\/\/en.wikipedia.org\/wiki\/Alnus_rubra<\/a><\/p>\n\n\n\n<p>Red alder is found in moist places, along streams, lake shores, and low lying areas of the landscape where water collects. It&#8217;s also a &#8216;pioneer species&#8217;, meaning that it&#8217;s often the first one to colonize a recently cleared area, e.g. after a landslide or forest fire. <\/p>\n\n\n\n<p>Red alder is significant in that it engages in a symbiotic relationships with nitrogen-fixing actinomycete bacteria of the genus <em>Frankia<\/em>. The bacteria take up residence in the root system of red alder. The tree provides their bacterial symbiont with a protected habitat and carbohydrates for food, while the bacteria provide their host tree with nitrogen in the form of ammonia. Useful forms of nitrogen (ammonia and nitrate) are typically in short supply in the soil, so this symbiotic relationship gives red alder an adaptive advantage. Eventually, the nitrogen fixed by <em>Frankia<\/em> and used by the red alder dissipates to the rest of the ecosystem. While most trees are careful about breaking down proteins and retrieving all the precious amino acids and other nitrogen-containing compounds from their leaves before dropping them in the autumn, red alder can afford to drop its leaves while they&#8217;re still green and rich in nitrogen-containing compounds. As the alder leaves are degraded by microbes and fungi in the soil, ammonia and nitrates are released and made available to other plants in the habitat. In that sense, red alder (with assistance from <em>Frankia<\/em>) helps the growth of other plants around it.    <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"768\" height=\"1024\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Alnus-incana-nodule_Jesse-Bellemare-768x1024.jpg\" alt=\"\" class=\"wp-image-267\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Alnus-incana-nodule_Jesse-Bellemare-768x1024.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Alnus-incana-nodule_Jesse-Bellemare-225x300.jpg 225w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Alnus-incana-nodule_Jesse-Bellemare-788x1051.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Alnus-incana-nodule_Jesse-Bellemare.jpg 900w\" sizes=\"auto, (max-width: 768px) 100vw, 768px\" \/><figcaption>Root nodules on a related alder species (<em>Alnus incana<\/em>). Symbiotic actinomycetes, <em>Frankia<\/em>, reside in these nodules. <em>Alnus rubra<\/em>, red alder, makes similar root nodules. Photo credit: Jesse Bellemere via Twitter<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"768\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall-1024x768.jpg\" alt=\"\" class=\"wp-image-613\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall-1024x768.jpg 1024w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall-300x225.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall-768x576.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall-1536x1152.jpg 1536w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall-788x591.jpg 788w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/07\/alder-leaves-green-in-fall.jpg 1700w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Red alder leaves, dropped while still green in the fall. Note the bigleaf maple (<a href=\"https:\/\/wordpress.viu.ca\/biol223\/acer-macrophyllum\/\"><em>Acer macrophyllum<\/em><\/a>) leaves which were yellow-brown when dropped, reflecting the fact that proteins have been degraded and nitrogen-rich molecules retracted into the trunk of the bigleaf maple tree for use again next year.  <\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"667\" height=\"1000\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-grey-trunks_smaller.jpg\" alt=\"\" class=\"wp-image-259\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-grey-trunks_smaller.jpg 667w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-grey-trunks_smaller-200x300.jpg 200w\" sizes=\"auto, (max-width: 667px) 100vw, 667px\" \/><figcaption>While walking through the woods, pale grey trunks with whitish splotches of lichen growth alerts you to the presence of red alder. Photo credit: Staffan Lindgren<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"667\" height=\"1000\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-grey-trunk.jpg\" alt=\"\" class=\"wp-image-258\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-grey-trunk.jpg 667w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-grey-trunk-200x300.jpg 200w\" sizes=\"auto, (max-width: 667px) 100vw, 667px\" \/><figcaption>A close up of a red alder trunk, showing the light grey color of the bark, and the white patches of lichens growing on the surface. The short horizontal raised marks on the bark are called &#8216;lenticels&#8217;. Lenticels are porous areas of the bark, allowing oxygen to diffuse into, and carbon dioxide to diffuse out of the living, respiring tissue under the bark. Photo credit: Staffan Lindgren<\/figcaption><\/figure>\n\n\n\n<p>Why is the species called &#8216;red&#8217; alder? The name comes from the color of the freshly cut wood, as shown in the photo below. The red pigment is especially intense in the inner bark, and has traditional uses by First Nations people in our region to, e.g., dye red cedar fibers used for weaving.  Alder wood is considered one of the best for smoking salmon. <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"400\" height=\"300\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/red-alder-wood_Willamette-Biology-via-Flickr.jpg\" alt=\"\" class=\"wp-image-266\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/red-alder-wood_Willamette-Biology-via-Flickr.jpg 400w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/red-alder-wood_Willamette-Biology-via-Flickr-300x225.jpg 300w\" sizes=\"auto, (max-width: 400px) 100vw, 400px\" \/><figcaption>The freshly cut wood of red alder is, as the name suggests, quite red. The colour pales as the wood dries and ages. Photo credit: Willamette Biology via Flickr<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"667\" height=\"1000\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-foliage_smaller.jpg\" alt=\"\" class=\"wp-image-257\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-foliage_smaller.jpg 667w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-foliage_smaller-200x300.jpg 200w\" sizes=\"auto, (max-width: 667px) 100vw, 667px\" \/><figcaption>The foliage of red alder appears quite shiny and dark green. Photo credit: Staffan Lindgren<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1000\" height=\"667\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-twig_1.jpg\" alt=\"\" class=\"wp-image-261\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-twig_1.jpg 1000w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-twig_1-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-twig_1-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-twig_1-788x526.jpg 788w\" sizes=\"auto, (max-width: 1000px) 100vw, 1000px\" \/><figcaption>Twig of red alder, note the alternate arrangement of leaves along the stem. Photo credit: Staffan Lindgren<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1000\" height=\"666\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-leaves.jpg\" alt=\"\" class=\"wp-image-260\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-leaves.jpg 1000w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-leaves-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-leaves-768x511.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-leaves-788x525.jpg 788w\" sizes=\"auto, (max-width: 1000px) 100vw, 1000px\" \/><figcaption>Note the characteristic venation of the red alder leaf, as well as the toothed edge. Notice that the edge is wavy, and also finely toothed. If you turned the leaf over, you&#8217;d notice that the edge looked as it had been very carefully rolled under. Photo credit: Staffan Lindgren<\/figcaption><\/figure>\n\n\n\n<p>Red alder is a flowering plant, but its flowers are not the typical shape that we tend to think. The cone in the photo below is an old female inflorescence, a cylindrical spike of many female flowers. And the longer hanging spike is the male inflorescence. <\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1000\" height=\"615\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-catkin-cone.jpg\" alt=\"\" class=\"wp-image-255\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-catkin-cone.jpg 1000w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-catkin-cone-300x185.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-catkin-cone-768x472.jpg 768w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/Red-alder-catkin-cone-788x485.jpg 788w\" sizes=\"auto, (max-width: 1000px) 100vw, 1000px\" \/><figcaption>Male and female flowers of red alder. The female inflorescence (cluster of several flowers) is the egg-shaped cone. The male inflorescence is the longer hanging spike (&#8216;catkin&#8217;). Both of these are old flowers remaining on the tree after reproduction is concluded. Photo credit: Staffan Lindgren<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"400\" height=\"254\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/red-alder-female-green-born1945-Flickr.jpg\" alt=\"\" class=\"wp-image-264\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/red-alder-female-green-born1945-Flickr.jpg 400w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/red-alder-female-green-born1945-Flickr-300x191.jpg 300w\" sizes=\"auto, (max-width: 400px) 100vw, 400px\" \/><figcaption>Immature green female red alder cones in spring. Note the leaf edge on the right &#8211; the edge looks rolled under.  Photo credit: born1945 via Flickr<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"400\" height=\"400\" src=\"http:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/male-catkin-andrew-reding-chestnut-backed-chickadee.jpg\" alt=\"\" class=\"wp-image-263\" srcset=\"https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/male-catkin-andrew-reding-chestnut-backed-chickadee.jpg 400w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/male-catkin-andrew-reding-chestnut-backed-chickadee-300x300.jpg 300w, https:\/\/wordpress.viu.ca\/biol223\/files\/2020\/06\/male-catkin-andrew-reding-chestnut-backed-chickadee-150x150.jpg 150w\" sizes=\"auto, (max-width: 400px) 100vw, 400px\" \/><figcaption>Male red alder catkin in spring, feasted upon by a chestnut backed chickadee. Photo credit: Andrew Reding via Flickr<\/figcaption><\/figure>\n","protected":false},"excerpt":{"rendered":"<p>Family: Betulaceae Common name: Red alder E-flora BC: https:\/\/linnet.geog.ubc.ca\/Atlas\/Atlas.aspx?sciname=Alnus%20rubra Wikipedia: https:\/\/en.wikipedia.org\/wiki\/Alnus_rubra Red alder is found in moist places, along streams, lake shores, and low lying areas of the landscape where [&hellip;]<\/p>\n","protected":false},"author":2649,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_monsterinsights_skip_tracking":false,"_monsterinsights_sitenote_active":false,"_monsterinsights_sitenote_note":"","_monsterinsights_sitenote_category":0,"footnotes":""},"class_list":["post-252","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages\/252","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/users\/2649"}],"replies":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/comments?post=252"}],"version-history":[{"count":8,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages\/252\/revisions"}],"predecessor-version":[{"id":718,"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/pages\/252\/revisions\/718"}],"wp:attachment":[{"href":"https:\/\/wordpress.viu.ca\/biol223\/wp-json\/wp\/v2\/media?parent=252"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}