{"id":2248,"date":"2019-10-25T00:08:53","date_gmt":"2019-10-25T00:08:53","guid":{"rendered":"http:\/\/wordpress.viu.ca\/biol325\/?p=2248"},"modified":"2019-12-04T19:11:21","modified_gmt":"2019-12-04T19:11:21","slug":"shakespeares-greatest-tragedy-the-invasive-european-starling","status":"publish","type":"post","link":"https:\/\/wordpress.viu.ca\/biol325\/2019\/10\/25\/shakespeares-greatest-tragedy-the-invasive-european-starling\/","title":{"rendered":"Shakespeare&#8217;s Greatest Tragedy: The Invasive European Starling"},"content":{"rendered":"\n<p class=\"wp-block-paragraph\">The year\nwas 1890, Europeans <em>flocked<\/em>\nto the big apple for a chance to begin a better life in the new world. Bringing\nwith them shoes, hats, smallpox,\nand the European Starling. <\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter is-resized\"><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-1-2-1024x168.png\" alt=\"\" class=\"wp-image-2357\" width=\"752\" height=\"123\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-1-2-1024x168.png 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-1-2-300x49.png 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-1-2-768x126.png 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-1-2.png 2045w\" sizes=\"auto, (max-width: 752px) 100vw, 752px\" \/><\/figure><\/div>\n\n\n\n<p class=\"wp-block-paragraph\">As the legend has it, an avid Shakespeare fan, and lover of birds, by the name of Eugene Schieffelin introduced 60 Starlings to Central Park New York City with the dignified intent of introducing all species of bird mentioned in Shakespeare&#8217;s writings  (<a href=\"https:\/\/www.smithsonianmag.com\/science-nature\/the-invasive-species-we-can-blame-on-shakespeare-95506437\/\">Zielinski 2011<\/a>). The Nightingales and the Skylarks he had previously released had all died off, so he could only cross his fingers and hope that the Starlings would breed (<a href=\"https:\/\/www.smithsonianmag.com\/science-nature\/the-invasive-species-we-can-blame-on-shakespeare-95506437\/\">Zielinski 2011<\/a>)! And indeed, breed they did. Today, 129 years later, North America is home to over 200 million Starling ranging geographically from Alaska all the way down to central America (<a href=\"https:\/\/birdsna.org\/Species-Account\/bna\/species\/eursta\/introduction\">Cabe 1993<\/a>). Thanks Eugene!<\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter is-resized\"><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Distn.png\" alt=\"\" class=\"wp-image-2249\" width=\"444\" height=\"525\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Distn.png 541w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Distn-254x300.png 254w\" sizes=\"auto, (max-width: 444px) 100vw, 444px\" \/><figcaption> Figure 1: The North American geographical distribution of the European Starling. There are actually less European Starling in Europe than in North America at this point as their European numbers have been declining due to fewer grassland invertebrates being available as a food source (<a href=\"https:\/\/onlinelibrary.wiley.com\/doi\/full\/10.1111\/ele.12387?casa_token=Yynavit6yeYAAAAA%3A22kSXuDPf_laUBa9BrWctIicILUw7yHpjukWSzrzCanORavN5VNlnrF435X0teF2NYT9QpBwJST8\">Inger et al. 2015<\/a>).<\/figcaption><\/figure><\/div>\n\n\n\n<hr class=\"wp-block-separator is-style-wide\" \/>\n\n\n\n<h2 class=\"wp-block-heading\">Meet The European Starling<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">Starling\u2019s are moderate sized passerine bird that is very similar in size to an American Robin; the average European Starling has a 16 inch wing span, is about 8 and a half inches long, and weighs in at 82 grams (<a href=\"https:\/\/www.amazon.ca\/Sibley-Field-Guide-Western-America\/dp\/0307957926\/ref=pd_sbs_14_2\/142-6277658-8595347?_encoding=UTF8&amp;pd_rd_i=0307957926&amp;pd_rd_r=0b782b79-2436-47f2-b8e5-d511668bd3ff&amp;pd_rd_w=G0UZT&amp;pd_rd_wg=FIsxh&amp;pf_rd_p=0602d3b5-e536-4dc4-9e55-dd650b3d14d4&amp;pf_rd_r=8W58X6Q1WYMTCEZ199AY&amp;psc=1&amp;refRID=8W58X6Q1WYMTCEZ199AY\">Sibley 2000<\/a>). If you are looking to spot a European Starling the first thing you need to know is that they have differing plumages throughout their life.<\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"alignright is-resized\"><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-3.png\" alt=\"\" class=\"wp-image-2347\" width=\"325\" height=\"253\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-3.png 698w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-3-300x234.png 300w\" sizes=\"auto, (max-width: 325px) 100vw, 325px\" \/><\/figure><\/div>\n\n\n\n<p class=\"wp-block-paragraph\">In breeding season (between spring and summer) the European Starling is very striking. A mysterious colour palette of iridescent deep purples, blues, and greens accompanied by a distinguishable yellow bill combine to give this bird a sinister beauty  \u2014 sinister because they are horribly invasive, and beauty because they are quite beautiful (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>; <a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">The Cornell Lab&#8230; 2019<\/a>). In non-breeding season the bill will usually be black but will turn yellow as breeding season grows near and the colours of the plumage are the same. However, you will be able to see cream coloured spots speckled about their body. This is because when their feathers are first grown they have cream tips (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">The Cornell Lab&#8230; 2019<\/a>). &nbsp;<\/p>\n\n\n\n<figure class=\"wp-block-image\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"251\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Comparison-1024x251.png\" alt=\"\" class=\"wp-image-2318\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Comparison-1024x251.png 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Comparison-300x73.png 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Comparison-768x188.png 768w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Figure 2: The differing plumage of European Starlings. Breeding adult (left), Non-breeding adult (middle), juvenile (right) (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">Photos by Dan Vickers &#8211; left, Ryan Schain &#8211; middle, and Walter Marcisz &#8211; right<\/a>)<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">By breeding season all these spots will be gone, but not by molting as you might have suspected. Instead, the spots disappear by simply being worn off &nbsp;(<a href=\"https:\/\/www.amazon.ca\/Sibley-Field-Guide-Western-America\/dp\/0307957926\/ref=pd_sbs_14_2\/142-6277658-8595347?_encoding=UTF8&amp;pd_rd_i=0307957926&amp;pd_rd_r=0b782b79-2436-47f2-b8e5-d511668bd3ff&amp;pd_rd_w=G0UZT&amp;pd_rd_wg=FIsxh&amp;pf_rd_p=0602d3b5-e536-4dc4-9e55-dd650b3d14d4&amp;pf_rd_r=8W58X6Q1WYMTCEZ199AY&amp;psc=1&amp;refRID=8W58X6Q1WYMTCEZ199AY\">Sibley 2016<\/a>). The feathers do not continue to wear down past the tips as the rest of the feather contains melanin (a dark pigment that is resistant to wear). Finally, in juvenile plumage the Starling will be a pale brown colour with no spots at all and a black bill (<a href=\"https:\/\/www.amazon.ca\/Sibley-Field-Guide-Western-America\/dp\/0307957926\/ref=pd_sbs_14_2\/142-6277658-8595347?_encoding=UTF8&amp;pd_rd_i=0307957926&amp;pd_rd_r=0b782b79-2436-47f2-b8e5-d511668bd3ff&amp;pd_rd_w=G0UZT&amp;pd_rd_wg=FIsxh&amp;pf_rd_p=0602d3b5-e536-4dc4-9e55-dd650b3d14d4&amp;pf_rd_r=8W58X6Q1WYMTCEZ199AY&amp;psc=1&amp;refRID=8W58X6Q1WYMTCEZ199AY\">Sibley 2016<\/a>).<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">For further identification traits it is helpful to know that they tend to fly in flocks \u2014 known as murmurations, but more on that below\u2014 and their flight pattern is a sustainable powered flight with very little gliding (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>). A wise Frenchmen once told me that the European Starling is easily identifiable by its short tail and its pointed wing tips (2019 advice from Eric Demers)\u2026 Moments later he confused a Raven for an Eagle, but that&#8217;s beside the point.<\/p>\n\n\n\n<figure class=\"wp-block-image\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"545\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Black-bird-starling-comp-1024x545.png\" alt=\"\" class=\"wp-image-2319\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Black-bird-starling-comp-1024x545.png 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Black-bird-starling-comp-300x160.png 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Black-bird-starling-comp-768x409.png 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Black-bird-starling-comp.png 1147w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption>Figure 4: From a distance the European Starling could be confused with other bird species such as the red winged blackbird (left). However, note the significant difference in wing and tail shape between the two birds (Photos by <a href=\"https:\/\/www.flickr.com\/photos\/neilscot\/7436137312\/\">Ian Neilson &#8211; left<\/a>, and <a href=\"https:\/\/flickr.com\/photos\/pware\/6436286829\">Patricia Ware &#8211; right<\/a>)<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">European Starling\u2019s don\u2019t like to eat alone, they primarily feed on insects and prefer to forage on the ground but may also feed in flight (<a href=\"https:\/\/www.audubon.org\/field-guide\/bird\/european-starling\">Audubon 2019<\/a>). Skull and jaw muscle adaptations have been noted to allow the bird to spring its mouth open to help with probing and feeding. Upon inserting their bill into the soil the European Starling&#8217;s strong jaw muscles allow it to spring open and widen the hole (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">The Cornell Lab&#8230; 2019<\/a>). Furthermore, their narrow skull makes it possible for the eye to move forward allowing the European Starling to peer into the hole and identify any food items (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>). <\/p>\n\n\n\n<p class=\"wp-block-paragraph\">While the European Starling seems to prefer feeding on insects, they are very versatile and adaptive feeders; a skill that has no doubt helped them become so established world wide (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016;<\/a> <a href=\"https:\/\/www.audubon.org\/field-guide\/bird\/european-starling\">Audubon 2019;<\/a> <a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">The Cornell Lab&#8230; 2019<\/a>). They have been documented to eat fruits, insects, spiders, snails, seeds, and even frogs and lizards  (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a> ). Considering just how far and wide this bird has travelled it should come as no surprise that they eat such a cultured diet. <\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Because of this diet the European Starling has been referred to as a pest by many. These guys might just come along and dig up your newly planted seeds, or maybe they&#8217;ve decided to eat your grapes right off the vine. Either way, when a large group of Starling come around your crops you could find yourself faced with a lot of damage. A 1996 study estimated that European Starlings caused 800 million dollars of agricultural damage per year in the United States alone (<a href=\"https:\/\/bioone.org\/journals\/BioScience\/volume-50\/issue-1\/0006-3568(2000)050[0053:EAECON]2.3.CO;2\/Environmental-and-Economic-Costs-of-Nonindigenous-Species-in-the-United\/10.1641\/0006-3568(2000)050[0053:EAECON]2.3.CO;2.full?casa_token=d3TUs8p3CBMAAAAA:d6jbj8cZOZF-aZgwruaQfL_Y2r6wElOLR27glXeL65HFsQ4hJ1UIwUkNsazg_POxd_0t1Ljw\">Pimental et al. 2000<\/a>).<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">On top of being an agricultural pest, the European Starling is also a pest to native bird species. European Starlings tend to be particular about their nesting sites, preferring to nest in pre-established cavities (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">The Cornell Lab&#8230; 2019<\/a>). Well what happens if they like the nesting nook of another bird?  Unfortunately, they have been known to evict many native species from their nesting sites, including: Wood ducks, Woodpeckers, Screech Owls, Kestrel, and undoubtedly many more (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>). European Starlings even tend to torment their kin, up to one third of a European Starling\u2019s first broods are killed by fellow Starlings dumping eggs out of the nest (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>).<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">While we deem the Starling to be a bit of a pest it seems that they enjoy being around us as they are most often found in disturbed habitat such as suburbs, fields, and farms and are rarely seen in extensive forests. Some European Starling may migrate for a warmer climate in the winter, yet others choose not too (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>). As can be seen in <em>figure 1<\/em>, they are found year-round in almost all of North America.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">The European Starling has been known to dabble in vocal mimicry and is classified as an intermediate mimic meaning it does mimic but predominately uses its own vocalizations (<a href=\"https:\/\/www.jstor.org\/stable\/4534373?casa_token=ePLXy1vdS6AAAAAA:djTqzKO0z1-bQbkx7wvhhnnlJyN7Y8F7PRYD3op9D4vl6oLdoUXWFwwwpR813E7IzEyc3-RnHa5ETcwwj52e3u5140EHFogcilqbSnZtr1AQwj_NpQ&amp;seq=1#metadata_info_tab_contents\">Hindmarsh 1984<\/a>). While the European Starling can mimic, their repertoire is limited to songs and sounds that are simple in structure but they can still learn dozens of different songs (<a href=\"https:\/\/www.jstor.org\/stable\/4534373?casa_token=ePLXy1vdS6AAAAAA:djTqzKO0z1-bQbkx7wvhhnnlJyN7Y8F7PRYD3op9D4vl6oLdoUXWFwwwpR813E7IzEyc3-RnHa5ETcwwj52e3u5140EHFogcilqbSnZtr1AQwj_NpQ&amp;seq=1#metadata_info_tab_contents\">Hindmarsh 1984<\/a>). As with many mimics, these mimicked vocalizations are used as a breeding strategy by males for attracting a female. However, both males and females can mimic and are known to sing outside of breeding season (<a href=\"http:\/\/www.sialis.org\/starlingbio.htm\">Sialis 2016<\/a>).<\/p>\n\n\n\n<figure class=\"wp-block-audio\"><audio controls src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Starling-Song.mp3\"><\/audio><figcaption>Some standard sounds you may hear in a European Starling Song. Mostly whistles and mechanical sounding clicks (Audio from <a href=\"https:\/\/identify.whatbird.com\/obj\/160\/_\/European_Starling.aspx\">Whatbird.com<\/a>)<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-embed-youtube wp-block-embed is-type-video is-provider-youtube wp-embed-aspect-4-3 wp-has-aspect-ratio\"><div class=\"wp-block-embed__wrapper\">\n<iframe loading=\"lazy\" title=\"Starling mimics hawk\" width=\"1180\" height=\"885\" src=\"https:\/\/www.youtube.com\/embed\/oU4Lbnf5Fm8?start=15&#038;feature=oembed\" frameborder=\"0\" allow=\"accelerometer; autoplay; encrypted-media; gyroscope; picture-in-picture\" allowfullscreen><\/iframe>\n<\/div><figcaption>An awesome video showing off the European Starlings mimicking skills with a recognizable <s>Bald Eagle<\/s> Red-tailed Hawk call! (Video by <a href=\"https:\/\/www.youtube.com\/watch?v=oU4Lbnf5Fm8\">ShadeTree1432<\/a>)<\/figcaption><\/figure>\n\n\n\n<hr class=\"wp-block-separator is-style-wide\" \/>\n\n\n\n<h2 class=\"wp-block-heading\">A Murmuration Of Starling<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">Flocks of birds all have bizarre names\u2026 I encourage you to check out <a rel=\"noreferrer noopener\" aria-label=\" (opens in a new tab)\" href=\"https:\/\/www.thespruce.com\/flock-names-of-groups-of-birds-386827\" target=\"_blank\">this list<\/a> if you are looking for some quality procrastination! A flock of Starling is known as a murmuration. Murmurations can be breath taking and fascinating to observe. Like a perfect sunset it\u2019s almost impossible for a photo or video to do this phenomenon justice. The movement that is sometimes displayed during a murmuration is simply astounding, it\u2019s almost as if the birds are one hive-mind entity. The exact reason why Starling\u2019s participate in murmurations isn\u2019t fully understood, but there are some leading hypotheses \u2014 before we go any further let&#8217;s take a look at a murmuration!<\/p>\n\n\n\n<figure class=\"wp-block-embed-youtube wp-block-embed is-type-video is-provider-youtube wp-embed-aspect-16-9 wp-has-aspect-ratio\"><div class=\"wp-block-embed__wrapper\">\n<iframe loading=\"lazy\" title=\"Flight of the Starlings: Watch This Eerie but Beautiful Phenomenon | Short Film Showcase\" width=\"1180\" height=\"664\" src=\"https:\/\/www.youtube.com\/embed\/V4f_1_r80RY?feature=oembed\" frameborder=\"0\" allow=\"accelerometer; autoplay; encrypted-media; gyroscope; picture-in-picture\" allowfullscreen><\/iframe>\n<\/div><figcaption><a href=\"https:\/\/www.youtube.com\/watch?v=V4f_1_r80RY\">National Geographic<\/a> footage of a murmuration of European Starling shot in the Netherlands.<br>Obligatory &#8220;that&#8217;s pretty neat!&#8221; <\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Have you ever heard the expression \u201cYou don\u2019t have to outrun a bear. You only have to outrun your slowest friend\u201d? Well, that expression just might hold true when it comes to starlings. Some researchers propose European Starlings may use murmurations to &#8216;dilute&#8217; themselves into a flock and lower their chances of being snatched up by avian predators such as falcons and hawks (<a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0179277\">Goodenough et al. 2017<\/a>). Okay so maybe this is the reason large groups of European Starling come together, but you might be wondering why they act out such eerie undulations. Of course, researcher\u2019s have a hypothesis for that \u2014 the \u201cselfish herd\u201d theory. <\/p>\n\n\n\n<p class=\"wp-block-paragraph\">The selfish herd theory suggests that the individual Starlings may be trying to avoid the edge of the murmuration, where they are most susceptible to predation, by moving towards the centre (<a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0179277\">Goodenough et al. 2017<\/a>). Since all birds are constantly trying to &#8216;out-run-their-friend&#8217; to the centre it results in a dynamic centre that is constantly shifting and moving around.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">There\u2019s also strong reason to believe that the murmurations might exist to confuse predators (<a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347211002867\">Procaccini et al. 2011<\/a>; <a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0179277\">Goodenough et al. 2017<\/a>). The constant wave-like motions of the flock may be interfering with a predator\u2019s ability to lock onto a target. One study showed that these wave-like movements occurred much more frequently while under predation and that the waves propagated away from the predator. In this sense it\u2019s almost as if the predator is the source of the wave (<a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347211002867\">Procaccini et al. 2011<\/a>).<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">The question scientists can\u2019t help but ask is \u201chow exactly are these birds communicating and moving in unison?\u201d An interesting study by an Italian theoretical physicist focused on how the communication during murmuration can be explained by \u201cself-organized criticality\u201d which is the same principal behind how an avalanche occurs (<a href=\"https:\/\/www.pnas.org\/content\/107\/26\/11865\/\">Cavagna et al. 2010<\/a>)! In the simplest of terms self-organized criticality is a property of dynamic systems that have a critical point. Think of it like having a pile of sand and each additional grain of sand you drop on top causes a small avalanche of sand (<a href=\"https:\/\/everything.explained.today\/Self-organized_criticality\/\">Self-organized&#8230; 2018<\/a>). Now if we think of the dynamic system that is a starling murmuration, we can imagine that first bird to move in response to an environmental influence is the grain of sand that causes an avalanche. A starling moves and all nearby starlings follow suit causing a cascade of movement! On top of that mathematical analyses shows that all Starling\u2019s in a flock are influenced by one another no matter how far apart they are and no matter how big the flock is (<a href=\"https:\/\/www.pnas.org\/content\/107\/26\/11865\/\">Cavagna et al. 2010<\/a>). Still, these are birds we are talking about, not just grains of sand. It is quite frankly nothing short of incredible that they are able to achieve this level of connection.<\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"alignright is-resized\"><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-2.png\" alt=\"\" class=\"wp-image-2332\" width=\"257\" height=\"302\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-2.png 490w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/10\/Fun-fact-2-256x300.png 256w\" sizes=\"auto, (max-width: 257px) 100vw, 257px\" \/><\/figure><\/div>\n\n\n\n<p class=\"wp-block-paragraph\">Akin to a well-trained scouts\u2019 troop, the European Starling murmuration move as a well-organized group. The shape of the flock is almost always guaranteed to follow the same proportions and aspect ratio regardless of the size of the group (<a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347208001176\">Ballerini et al. 2008<\/a>). The murmurations are often thin and elongated and while this shape is well documented there is no consensus as to why this shape is adopted. It could be due to several factors including aerodynamics and even gravity making thin and wide flocks more favourable (<a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347208001176\">Ballerini et al. 2008<\/a>)! Murmuration dynamics have been studied from a physics, mathematics, and biological point of view for many years and continue to fascinate scientists from all fields with their extraordinary complexity.<\/p>\n\n\n\n<h2 class=\"wp-block-heading\">Tell me, where can I see this phenomenon?<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">Well I am glad you asked!<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">If you are looking to see a murmuration in person you will have the best chance by going to an area where European Starling are common such as fields and other disturbed areas. Seasonally European Starlings are known to have the largest murmurations in winter when more birds are in closer geographical proximity due to flocking to warmer climates \u2014 however, I have found that downtown Nanaimo during the summer months can be a great place to witness some awesome murmurations! <\/p>\n\n\n\n<p class=\"wp-block-paragraph\">The European Starling is a polarizing bird. Striking in image and mesmerizing in murmuration, yet they are an invasive species that has impacts on native ecosystems. Fortunately, they haven&#8217;t appeared to contribute the declines of most native bird populations so don\u2019t feel too guilty next time you are kicking back to &#8216;chillax&#8217; and enjoy a murmuration (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/id\">The Cornell Lab&#8230; 2019<\/a>). <\/p>\n\n\n\n<div style=\"height:143px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p class=\"wp-block-paragraph\">All about birds [Internet]. 2019. The cornell lab of ornithology: [cited 2019 oct 24]. Available from: <a href=\"https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/lifehistory.\">https:\/\/www.allaboutbirds.org\/guide\/European_Starling\/lifehistory.<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">All about starlings [Internet]. 2016. Sialis: [2019 oct 24]. Available from:  <a href=\"http:\/\/www.sialis.org\/starlingbio.htm.\">http:\/\/www.sialis.org\/starlingbio.htm.<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Ballerini M, Cabibbo N, Candelier R, Cavagna A, Cisbani E, Giardina I, Orlandi A, Parisi G, Procaccini A, Viale M, et al. 2008. Empirical investigation of starling flocks: a benchmark study in collective animal behaviour. Anim Behav. 76(1): 201\u2013215. Available from: <a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347208001176\">https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347208001176<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Cabe PR. 1993. European Starling (Sturnus vulgaris). The Cornell Laboratory of Ornithology. Available from: <a href=\"https:\/\/birdsna.org\/Species-Account\/bna\/species\/eursta\/introduction.\">https:\/\/birdsna.org\/Species-Account\/bna\/species\/eursta\/introduction.<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Cavagna A, Cimarelli A, Giardina I, Parisi G, Santagati R, Stefanini F, Viale M. 2010. Scale-free correlations in starling flocks. Proc Natl Acad Sci U S A. 107(26): 11865\u201311870. Available from: <a href=\"https:\/\/www.pnas.org\/content\/107\/26\/11865\/\">https:\/\/www.pnas.org\/content\/107\/26\/11865\/<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">European Starling [Internet]. 2019. Audubon; [cited 2019 oct 24]. Available from: <a href=\"https:\/\/www.audubon.org\/field-guide\/bird\/european-starling\">https:\/\/www.audubon.org\/field-guide\/bird\/european-starling<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Goodenough AE, Little N, Carpenter WS, Hart AG. 2017. Birds of a feather flock together: Insights into starling murmuration behaviour revealed using citizen science. PLoS One. 12(6): 1\u201318. Available from: <a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0179277\">https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0179277<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Hindmarsh AM. 1984. Vocal mimicry in starlings. Behaviour. 90(4), 302-324. Available from\u202f: <a href=\"https:\/\/www.jstor.org\/stable\/4534373?seq=1#metadata_info_tab_contents\">https:\/\/www.jstor.org\/stable\/4534373?seq=1#metadata_info_tab_contents<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Inger R, Gregory R, Duffy JP, Stott I, Vo\u0159\u00ed\u0161ek P, Gaston KJ. 2015. Common European birds are declining rapidly while less abundant species\u2019 numbers are rising. Ecol Lett. 18(1): 28\u201336. Available from: <a href=\"https:\/\/onlinelibrary.wiley.com\/doi\/abs\/10.1111\/ele.12387\">https:\/\/onlinelibrary.wiley.com\/doi\/abs\/10.1111\/ele.12387<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Pimental D, Lach L, Zuniga R, Morrison D. 2000. Environmental and economic costs of nonindigenous species in the united states. Bioscience. 50(1): 53. Available from: <a href=\"https:\/\/bioone.org\/journals\/BioScience\/volume-50\/issue-1\/0006-3568(2000)050[0053:EAECON]2.3.CO;2\/Environmental-and-Economic-Costs-of-Nonindigenous-Species-in-the-United\/10.1641\/0006-3568(2000)050[0053:EAECON]2.3.CO;2.short\">https:\/\/bioone.org\/journals\/BioScience\/volume-50\/issue-1\/0006-3568(2000)050[0053:EAECON]2.3.CO;2\/Environmental-and-Economic-Costs-of-Nonindigenous-Species-in-the-United\/10.1641\/0006-3568(2000)050[0053:EAECON]2.3.CO;2.short<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Procaccini A, Orlandi A, Cavagna A, Giardina I, Zoratto F, Santucci D, Chiarotti F, Hemelrijk CK, Alleva E, Parisi G, et al. 2011. Propagating waves in starling, Sturnus vulgaris, flocks under predation. Anim Behav. 82(4): 759\u2013765. Available from: <a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347211002867\">https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347211002867<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Self-organized criticality explained [Internet]. 2018. Everything explained today: [cited 2019 oct 24]. Available from: <a href=\"https:\/\/everything.explained.today\/Self-organized_criticality\/\">https:\/\/everything.explained.today\/Self-organized_criticality\/<\/a><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Sibley DA. 2016 Sibley Birds West. Second edition. New York (NY): Alfred A. Knopf. 477 p.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Zielinski S. [Internet]. 2011. The invasive species we can blame on shakespeare. Smithsonianmag.com [updated 2011 oct 4; cited 2019 oct 24]. Available from: <a href=\"https:\/\/www.smithsonianmag.com\/science-nature\/the-invasive-species-we-can-blame-on-shakespeare-95506437\/.\">https:\/\/www.smithsonianmag.com\/science-nature\/the-invasive-species-we-can-blame-on-shakespeare-95506437\/.<\/a><\/p>\n","protected":false},"excerpt":{"rendered":"<p>The year was 1890, Europeans flocked to the big apple for a chance to begin a better life in the new world. Bringing with them shoes, hats, smallpox, and the [&hellip;]<\/p>\n","protected":false},"author":1973,"featured_media":2414,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_crdt_document":"","footnotes":""},"categories":[1],"tags":[],"class_list":["post-2248","post","type-post","status-publish","format-standard","has-post-thumbnail","hentry","category-uncategorized"],"_links":{"self":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts\/2248","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/users\/1973"}],"replies":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/comments?post=2248"}],"version-history":[{"count":59,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts\/2248\/revisions"}],"predecessor-version":[{"id":3297,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts\/2248\/revisions\/3297"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/media\/2414"}],"wp:attachment":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/media?parent=2248"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/categories?post=2248"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/tags?post=2248"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}