{"id":2764,"date":"2019-11-02T19:11:55","date_gmt":"2019-11-02T19:11:55","guid":{"rendered":"http:\/\/wordpress.viu.ca\/biol325\/?p=2764"},"modified":"2019-12-02T19:05:40","modified_gmt":"2019-12-02T19:05:40","slug":"the-mallard","status":"publish","type":"post","link":"https:\/\/wordpress.viu.ca\/biol325\/2019\/11\/02\/the-mallard\/","title":{"rendered":"The Mallard"},"content":{"rendered":"\n<h3 class=\"wp-block-heading\"> Natural History <\/h3>\n\n\n\n<p class=\"wp-block-paragraph\"> Mallards are one of the most abundant ducks in the world, and likely the most familiar to passesby. The male Mallard has an iridescent green head and a dark brown breast separated by a thin white collar around the neck. Males in non-breeding plumage lose the identifying colours of their feathers but can still be identified by their bill colour. Females have orange and brown bills and mottled brown feathers. Both sexes can be identified by their speculum, a patch of feathers on the wing that\u2019s visible during flight, which is iridescent blue with a white border (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/Mallard\/id\">Allaboutbirds.org 2017<\/a>). <\/p>\n\n\n\n<figure class=\"wp-block-gallery columns-1 is-cropped wp-block-gallery-1 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"800\" height=\"532\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/Drey-and-Hen.jpg\" alt=\"\" data-id=\"2765\" class=\"wp-image-2765\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/Drey-and-Hen.jpg 800w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/Drey-and-Hen-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/Drey-and-Hen-768x511.jpg 768w\" sizes=\"auto, (max-width: 800px) 100vw, 800px\" \/><figcaption class=\"blocks-gallery-item__caption\"> <em>Female (left) and male (right) Mallard<\/em><br><em>Photo: <\/em><a href=\"https:\/\/www1.nyc.gov\/site\/wildlifenyc\/animals\/mallards.page\">WildlifeNYC<\/a> <\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<figure class=\"wp-block-gallery alignright columns-1 is-cropped wp-block-gallery-2 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"612\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/f548af70f1b767bf912f002b27b4168e-1024x612.jpg\" alt=\"\" data-id=\"2766\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/?attachment_id=2766\" class=\"wp-image-2766\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/f548af70f1b767bf912f002b27b4168e-1024x612.jpg 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/f548af70f1b767bf912f002b27b4168e-300x179.jpg 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/f548af70f1b767bf912f002b27b4168e-768x459.jpg 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/f548af70f1b767bf912f002b27b4168e.jpg 1600w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"blocks-gallery-item__caption\"> <em>Male Mexican Duck<\/em><br><em>Photo: <\/em><a href=\"http:\/\/200birds.blogspot.com\/2012\/02\/mexican-duck-in-utah.html\">Ryan O&#8217;Donnel<\/a> <\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<figure class=\"wp-block-gallery alignright columns-1 is-cropped wp-block-gallery-3 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"569\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/2787-duck-hybrid-20160407-1024x569.jpg\" alt=\"\" data-id=\"2767\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/?attachment_id=2767\" class=\"wp-image-2767\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/2787-duck-hybrid-20160407-1024x569.jpg 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/2787-duck-hybrid-20160407-300x167.jpg 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/2787-duck-hybrid-20160407-768x427.jpg 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/2787-duck-hybrid-20160407.jpg 1080w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"blocks-gallery-item__caption\"> <em>Male Mallard X American Black Duck hybrid<\/em><br><em>Photo: <\/em><a href=\"http:\/\/birdhybrids.blogspot.com\/2014\/04\/american-black-duck-x-mallard.html\">Gordon Johnston<\/a> <\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Mexican Ducks are thought to be a subspecies of Mallard, but genetic evidence indicates they are more closely related to the American Black Duck. Instead of a pronounced difference between males and females, both genders are mottled brown like the northern female. Mallards will also hybridize with other duck species, like the American Black Duck, which creates a blend of colourations and patterns (<a href=\"https:\/\/neotropical.birds.cornell.edu\/Species-Account\/nb\/species\/mexduc\/overview\">Neotropical.birds.cornell.edu 2019<\/a>). <\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\n\nMallards are dabbling ducks, submerging the head and neck while upending to probe the mud for food. They have an omnivorous diet consisting of seeds, roots, and grasses, as well as insects and small aquatic vertebrates. Mallards will also graze on land for these foods (<a href=\"https:\/\/www.audubon.org\/field-guide\/bird\/mallard\">Audubon.org 2019<\/a>). They will also accept food from people at duck ponds and city parks. People commonly feed bread to wild ducks, which doesn\u2019t provide the variety of nutrients present in their natural diet. As a substitute for bread, consider feeding ducks peas, corn, grains, seeds, or lettuce instead (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/Mallard\/id\">Allaboutbirds.org 2017<\/a>).\n\n<\/p>\n\n\n\n<figure class=\"wp-block-gallery columns-1 is-cropped wp-block-gallery-4 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/SPR_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e-1024x683.png\" alt=\"\" data-id=\"3108\" data-full-url=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/SPR_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e.png\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/2019\/11\/02\/the-mallard\/spr_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e\/\" class=\"wp-image-3108\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/SPR_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e-1024x683.png 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/SPR_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e-300x200.png 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/SPR_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e-768x512.png 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/SPR_386584-what-to-feed-ducks1-5b4671d3c9e77c001a0b015e.png 1500w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><\/figure><\/li><\/ul><figcaption class=\"blocks-gallery-caption\"><em>What to feed (and not feed!) ducks \/ <\/em>Photo: <a href=\"https:\/\/www.thespruce.com\/is-feeding-ducks-bread-bad-386564\">Hugo Lin<\/a><\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-gallery alignleft columns-1 is-cropped wp-block-gallery-5 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"713\" height=\"500\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/male-mallard-dabbling_1366127388_1200.jpg\" alt=\"\" data-id=\"3101\" data-full-url=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/male-mallard-dabbling_1366127388_1200.jpg\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/2019\/11\/02\/the-mallard\/male-mallard-dabbling_1366127388_1200\/\" class=\"wp-image-3101\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/male-mallard-dabbling_1366127388_1200.jpg 713w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/male-mallard-dabbling_1366127388_1200-300x210.jpg 300w\" sizes=\"auto, (max-width: 713px) 100vw, 713px\" \/><figcaption class=\"blocks-gallery-item__caption\"><em>Male Mallard dabbling<\/em><br>Photo: <a href=\"http:\/\/www.pixmix.ca\/photos\/nature\/birds\/waterfowl\/dabblers\/male-mallard-dabbling_1366127388.jpg.php\">Karen VK<\/a><\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">\n\nThey are widespread and common in the northern hemisphere, particularly in America, Canada, and Europe. <a href=\"https:\/\/ebird.org\/map\/mallar3?env.minX=-178.203369424671&amp;env.minY=-54.969919623211&amp;env.maxX=179.99999999291&amp;env.maxY=80.275133602445\">Click here<\/a> to see an interactive map of Mallard distribution. Some Mallards that breed in northern Canada and Alaska will migrate to the southern United States to over-winter. Others do not migrate and can be found year-round in most of America and coastal British Colombia (<a href=\"https:\/\/www.audubon.org\/field-guide\/bird\/mallard\">Audubon.org 2019<\/a>). Mallards will live in most wetlands, both natural and artificial, from lush marshes to roadside ditches . Mallards are the most heavily hunted duck, estimating a third of all ducks shot in North America (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/Mallard\/id\">Allaboutbirds.org 2017<\/a>). The Mallard is not considered at risk, but the current population trend is decreasing (<a href=\"https:\/\/www.nationalgeographic.com\/animals\/birds\/m\/mallard\/\">Nationalgeographic.com 2019<\/a>).&nbsp;\n\n<\/p>\n\n\n\n<figure class=\"wp-block-gallery columns-1 is-cropped wp-block-gallery-6 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/mallard9-59511df05f9b58f0fc660403-1024x683.jpg\" alt=\"\" data-id=\"2768\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/?attachment_id=2768\" class=\"wp-image-2768\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/mallard9-59511df05f9b58f0fc660403-1024x683.jpg 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/mallard9-59511df05f9b58f0fc660403-300x200.jpg 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/mallard9-59511df05f9b58f0fc660403-768x512.jpg 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/mallard9-59511df05f9b58f0fc660403.jpg 1500w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"blocks-gallery-item__caption\"> <em>Mallard chicks<\/em><br><em> Photo: <\/em><a href=\"https:\/\/www.flickr.com\/photos\/goellnitz\/34655339051\/\">Jen Goellnitz<\/a> <\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\"> Mallards will migrate once they have formed a breeding pair. A variety of courtship behaviours are used to establish pairs, including head-bobbing (seen in the video below), swimming quickly with the head lowered to the surface of the water, and, in males, whistling while circling a female (<a href=\"https:\/\/www.allaboutbirds.org\/what-to-watch-for-duck-courtship-video\/\">Allaboutbirds.org 2015<\/a>) They nest on the ground or in protected cavities, and build nests out of sticks, grass and down (<a href=\"https:\/\/www.nationalgeographic.com\/animals\/birds\/m\/mallard\/\">Nationalgeographic.com 2019<\/a>). A nest will typically have between 1-13 cream coloured eggs, but on one occasion a female was observed associating with 29 ducklings. This extreme case be be the result of brood parasitism, when females will lay their eggs in another female&#8217;s nest (<a href=\"https:\/\/notornis.osnz.org.nz\/observation-female-mallard-anas-platyrhynchos-29-ducklings\">Seabrook-Davison, 2014<\/a>).<\/p>\n\n\n\n<figure class=\"wp-block-gallery alignright columns-1 is-cropped wp-block-gallery-7 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"809\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/5b296d581fb74.image_-1024x809.jpg\" alt=\"\" data-id=\"3099\" data-full-url=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/5b296d581fb74.image_.jpg\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/2019\/11\/02\/the-mallard\/5b296d581fb74-image_\/\" class=\"wp-image-3099\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/5b296d581fb74.image_-1024x809.jpg 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/5b296d581fb74.image_-300x237.jpg 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/5b296d581fb74.image_-768x607.jpg 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/5b296d581fb74.image_.jpg 1200w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"blocks-gallery-item__caption\"><em>Mallard nest<\/em><br>Photo: <a href=\"https:\/\/www.postbulletin.com\/life\/lifestyles\/nature-nut-duck-nests-capture-my-fancy\/article_4af3f578-73ff-11e8-8217-cfc3456aa94c.html\">Greg Munson<\/a><\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">Although conspecific brood parasitism (CBP) sounds awful (raising 29 ducklings is a hard job!) the negative effects on the parasitized female are typically quite low, and is fairly rare in Mallards. It&#8217;s estimated that CBP occurs in 0-10% of nests. Parastic females may choose a nest based on easy detectability and quality of the nest, or nesting female (<a href=\"http:\/\/citeseerx.ist.psu.edu\/viewdoc\/download?doi=10.1.1.670.8457&amp;rep=rep1&amp;type=pdf\">Kreisinger et al, 2010<\/a>). Parasitized nests tend to have fewer eggs than average, even after a second female has laid in them. This keeps the burden on the parasitized mother from growing too large, thus mitigating possible negative effects (<a href=\"https:\/\/academic.oup.com\/condor\/article-abstract\/93\/2\/340\/5185417\">Lokemoen, 1991<\/a>). When broods do become large, som consequences include decreased hatching success, increased incubation period, and increased risk of abandonment by the mother (<a href=\"http:\/\/citeseerx.ist.psu.edu\/viewdoc\/download?doi=10.1.1.670.8457&amp;rep=rep1&amp;type=pdf\">Kreisinger et al, 2010<\/a>). <\/p>\n\n\n\n<p class=\"wp-block-paragraph\"> Males will abandon the nest once the eggs have hatched, leaving the female to care for the offspring. Ducklings are ready to leave nest within 13 hours of hatching. The young know how to swim as soon as they hatch and can feed on their own while the mother leads them between the nest and feeding grounds  (<a href=\"https:\/\/www.allaboutbirds.org\/guide\/Mallard\/id\">Allaboutbirds.org 2017<\/a>). <\/p>\n\n\n\n<figure class=\"wp-block-embed-youtube wp-block-embed is-type-video is-provider-youtube wp-embed-aspect-16-9 wp-has-aspect-ratio\"><div class=\"wp-block-embed__wrapper\">\n<iframe loading=\"lazy\" title=\"Mallard Duck Mating Dance (2-12-2013)\" width=\"1180\" height=\"664\" src=\"https:\/\/www.youtube.com\/embed\/drK2bwzjY9c?feature=oembed\" frameborder=\"0\" allow=\"accelerometer; autoplay; encrypted-media; gyroscope; picture-in-picture\" allowfullscreen><\/iframe>\n<\/div><figcaption> <em>The head-bobbing courtship behaviour in Mallards. Video: <\/em><a href=\"https:\/\/www.youtube.com\/watch?v=drK2bwzjY9c\">Michigan&#8217;s Wildlife<\/a> <\/figcaption><\/figure>\n\n\n\n<h3 class=\"wp-block-heading\"> Mating Behaviour and Genital Morphology <\/h3>\n\n\n\n<p class=\"wp-block-paragraph\">\n\nCopulation in Mallards has been studied for many years and is still being studied today. Birds typically have simple genitalia, with males lacking external genitalia and females having simple vaginas. Waterfowl, including mallards, are the only birds that have a penis. A Mallard\u2019s penis is corkscrew-shaped and is coated in ridges and spines (<a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0000418\">Brennan et al, 2007<\/a>). The length of the phallus, the size of the testes, and the amount and size of ridges and spines all increase when forced copulation is common (<a href=\"https:\/\/www.jstor.org\/stable\/4089887?seq=1#page_scan_tab_contents\">Coker et al, 2002<\/a>). The phallus is stored in a sac inside the body until it is everted. Eversion of the phallus has been termed \u201cexplosive,\u201d with the phallus becoming fully erect within a third of a second (<a href=\"https:\/\/royalsocietypublishing.org\/doi\/10.1098\/rspb.2009.2139\">Brennan et al, 2009<\/a>). Because Mallards typically copulate while swimming, using an intromittent organ like a penis may ensure that sperm are delivered to the female and not lost in the water (<a href=\"https:\/\/www.jstor.org\/stable\/4089887?seq=1#page_scan_tab_contents\">Coker et al, 2002<\/a>).\n\n<\/p>\n\n\n\n<figure class=\"wp-block-gallery columns-1 is-cropped wp-block-gallery-8 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"760\" height=\"570\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/140506-genital_eed48a9306dcb896cfb0e13f8417c513.fit-760w.jpg\" alt=\"\" data-id=\"2769\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/?attachment_id=2769\" class=\"wp-image-2769\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/140506-genital_eed48a9306dcb896cfb0e13f8417c513.fit-760w.jpg 760w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/140506-genital_eed48a9306dcb896cfb0e13f8417c513.fit-760w-300x225.jpg 300w\" sizes=\"auto, (max-width: 760px) 100vw, 760px\" \/><figcaption class=\"blocks-gallery-item__caption\"> <em>Penis (top) and vagina (bottom) of the Mallard<\/em><br><em>Photo: <\/em><a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0000418\">Patricia Brennan<\/a> <\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">\n\nFemale Mallards have corkscrew-shaped vaginas that turn in the opposite direction as the penis. This creates a mechanical barrier that makes full eversion of the penis difficult (<a href=\"https:\/\/royalsocietypublishing.org\/doi\/10.1098\/rspb.2009.2139\">Brennan et al, 2009<\/a>). The vagina also has two different types of pouches. The first type of pouch is found near the cloaca, and likely act as \u201cdead-ends\u201d for the phallus, which can prevent sperm from reaching the egg. Sperm deposited in these pouches may even be ejected by the female. The second type of pouch is found closer to the utero-vaginal junction and is used for storing the sperm of successful males for fertilization. (<a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0000418\">Brennan et al, 2007<\/a>)\n\n<\/p>\n\n\n\n<figure class=\"wp-block-gallery alignright columns-1 is-cropped wp-block-gallery-9 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"768\" src=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/IMG_8488-1024x768.jpg\" alt=\"\" data-id=\"3104\" data-full-url=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/IMG_8488.jpg\" data-link=\"https:\/\/wordpress.viu.ca\/biol325\/2019\/11\/02\/the-mallard\/mallard-forced-copulation\/\" class=\"wp-image-3104\" srcset=\"https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/IMG_8488-1024x768.jpg 1024w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/IMG_8488-300x225.jpg 300w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/IMG_8488-768x576.jpg 768w, https:\/\/wordpress.viu.ca\/biol325\/files\/2019\/11\/IMG_8488.jpg 1280w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"blocks-gallery-item__caption\"><em>An unfortunate female<\/em> \/ Photo: <a href=\"http:\/\/therattlingcrow.blogspot.com\/2015\/03\/forced-copulation-in-mallards.html\">Africa Gomez<\/a><\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\">\n\n&nbsp;Forced copulation is common in waterfowl, which causes conflict between the sexes. Older literature describes this behaviour by males as \u201crape\u201d, but modern literature prefers to use \u201cforced copulation\u201d to avoid the use of the controversial term. Such behaviour is recognized by active pursuit, grasping, and overpowering of a female by a male. Many males have been observed pursuing a single female, and this aggressive behaviour may even lead to the death of the female. Pair copulations, in contrast, involve characteristic mating displays performed by both sexes (<a href=\"https:\/\/www.jstor.org\/stable\/4085762?seq=1#page_scan_tab_contents\">Burns et al, 1980<\/a>). It has been shown that high testosterone levels will contribute to higher rates of forced-copulation attempts (<a href=\"https:\/\/www.jstor.org\/stable\/4602131?seq=1#page_scan_tab_contents\">Ellen S. Davis, 2002a<\/a>).\n\n<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">\n\nThe antagonistic morphology present in Mallards likely coevolved as a method for females to control fertilization by unwanted males. (<a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0000418\">Brennan et al, 2007<\/a>) Pairs are established weeks or months before migration to the breeding grounds, and a female will not solicit extra-pair copulations. (<a href=\"https:\/\/royalsocietypublishing.org\/doi\/10.1098\/rspb.2009.2139\">Brennan et al, 2009<\/a>) Females may choose mates based on the ability to defend against other males. In a study by <a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347202930794\">Davis<\/a> (<a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347202930794\">2002b)<\/a>, females that paired to males with high testosterone, and presumably were more active in protection, were missing less feathers from forced copulation attempts. \n\n<\/p>\n\n\n\n<figure class=\"wp-block-embed-youtube wp-block-embed is-type-video is-provider-youtube wp-embed-aspect-16-9 wp-has-aspect-ratio\"><div class=\"wp-block-embed__wrapper\">\n<iframe loading=\"lazy\" title=\"True Facts About The Duck\" width=\"1180\" height=\"664\" src=\"https:\/\/www.youtube.com\/embed\/6k01DIVDJlY?feature=oembed\" frameborder=\"0\" allow=\"accelerometer; autoplay; encrypted-media; gyroscope; picture-in-picture\" allowfullscreen><\/iframe>\n<\/div><figcaption> <em>I hope this video makes you laugh as much as I did. Video: <\/em><a href=\"https:\/\/www.youtube.com\/watch?v=6k01DIVDJlY\">zefrank1<\/a> <\/figcaption><\/figure>\n\n\n\n<h3 class=\"wp-block-heading\">References<\/h3>\n\n\n\n<ol class=\"wp-block-list\"><li><a href=\"https:\/\/www.audubon.org\/field-guide\/bird\/mallard\">Audubon.org (2019) Mallard, National Audubon Society.<\/a><\/li><li><a href=\"https:\/\/www.allaboutbirds.org\/guide\/Mallard\/id\">Allaboutbirds.org (2017) Mallard, Cornell Lab of Ornithology.<\/a><\/li><li><a href=\"https:\/\/www.nationalgeographic.com\/animals\/birds\/m\/mallard\/\">Nationalgeographic.com (2019) Mallard, National Geographic Society.<\/a><\/li><li><a href=\"https:\/\/notornis.osnz.org.nz\/observation-female-mallard-anas-platyrhynchos-29-ducklings\">Seabrooke-Davison, M (2014) Observation of a female mallard (Anas platyrhynchos) with 29 ducklings. Notornis, 61: 51-53<\/a><\/li><li><a href=\"https:\/\/www.allaboutbirds.org\/what-to-watch-for-duck-courtship-video\/\">Allaboutbirds.org (2015) How to recognize duck courtship displays, Cornell Lab of Ornithology.<\/a><\/li><li><a href=\"https:\/\/neotropical.birds.cornell.edu\/Species-Account\/nb\/species\/mexduc\/overview\">Neotropical.birds.cornell.edu (2019) Mexican Duck, Cornell Lab of Ornithology.<\/a><\/li><li><a href=\"http:\/\/citeseerx.ist.psu.edu\/viewdoc\/download?doi=10.1.1.670.8457&amp;rep=rep1&amp;type=pdf\">Kreisinger, J., Munclinger, P., Javurkova, V., Albrecht, T. (2010) Analysis of Extra-Pair Paternity and Conspecific Brood Parasitism in Anas Platyrynchos using non-invasive techniques. J. Avian Biol., 41: 551-557.   doi: 10.1111\/j.1600-048X.2010.05002.x<\/a><\/li><li><a href=\"https:\/\/academic.oup.com\/condor\/article-abstract\/93\/2\/340\/5185417\">Lokemoen, J. T. (1991) Brood Parasitism among Waterfowl Nesting on Islands and Peninsulas in North Dakota. The Condor, 93(2): 340-345<\/a><\/li><li><a href=\"https:\/\/journals.plos.org\/plosone\/article?id=10.1371\/journal.pone.0000418\">Brennan, P. L. R., Prum, R. O., McCracken, K. G., Sorenson, M. D., Wilson, R. E., Birkhead, T. R. (2007) Coevolution of Male and Female Genital Morphology in Waterfowl. PloSONE, 2(5): e418  doi:10.1371\/journal.pone.0000418<\/a><\/li><li><a href=\"https:\/\/www.jstor.org\/stable\/4089887?seq=1#page_scan_tab_contents\">Coker, C. R., McKinney, F., Hays, H., Briggs, S. V., Cheng, K. M. (2002) Intromittent organ morphology and testis size in relation to mating system in waterfowl. Auk, 119(2): 403-413 <\/a><\/li><li><a href=\"https:\/\/royalsocietypublishing.org\/doi\/10.1098\/rspb.2009.2139\">Brennan, P. L. R., Clark, C. J., Prum, R. O. (2009)  Explosive eversion and functional morphology of the duck penis supports sexual conflict in waterfowl genitalia. Proc. R. Soc. B, 277: 1309-1314 doi:10.1098\/rspb.2009.2139<\/a><\/li><li><a href=\"https:\/\/www.jstor.org\/stable\/4085762?seq=1#page_scan_tab_contents\">Burns, J. T., Cheng, K. M., McKinney, F. (1980) Forced Copulation in Captive Mallards I. Fertilization of Eggs. Auk, 97: 875-879<\/a><\/li><li><a href=\"https:\/\/www.jstor.org\/stable\/4602131?seq=1#page_scan_tab_contents\">Ellen S. Davis (2002a) Male reproductive tactics in the mallard, Anas platyrhynchos: social and hormonal mechanisms. Behav Ecol Sociobiol, 52: 224-231 DOI 10. 1007\/s00265-002-05 1 3-z<\/a><\/li><li><a href=\"https:\/\/www.sciencedirect.com\/science\/article\/pii\/S0003347202930794\">Ellen S. Davis (2002b) Female choice and the benefits of mate guarding by male mallards. Animal Behaviour, 64: 619-628 doi:10.1006\/anbe.2002.3079<\/a><\/li><\/ol>\n","protected":false},"excerpt":{"rendered":"<p>Natural History Mallards are one of the most abundant ducks in the world, and likely the most familiar to passesby. The male Mallard has an iridescent green head and a [&hellip;]<\/p>\n","protected":false},"author":1903,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_crdt_document":"","footnotes":""},"categories":[1],"tags":[],"class_list":["post-2764","post","type-post","status-publish","format-standard","hentry","category-uncategorized"],"_links":{"self":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts\/2764","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/users\/1903"}],"replies":[{"embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/comments?post=2764"}],"version-history":[{"count":3,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts\/2764\/revisions"}],"predecessor-version":[{"id":3109,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/posts\/2764\/revisions\/3109"}],"wp:attachment":[{"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/media?parent=2764"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/categories?post=2764"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/wordpress.viu.ca\/biol325\/wp-json\/wp\/v2\/tags?post=2764"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}